In Canada, "the author of a work shall be the first owner of the copyright therein" (Copyright Act § 13(1)).
Also, "[w]here the author of a work was in the employment of some other person under a contract of service or apprenticeship and the work was made in the course of his employment by that person, the person by whom the author was employed shall, in the absence of any agreement to the contrary, be the first owner of the copyright" (Copyright Act § 13(3)).
Nothing in your description of the situation describes to me an employer/employee relationship, so in my opinion, § 13(3) is not implicated.
Even if there were an agreement or contract to transfer copyright to an LLC or assign it to the other individuals, absent an actual writen assignment, the original owner still owns the copyright. "[N]o assignment or grant is valid unless it is in writing signed by the owner of the right..." (Copyright Act § 13(4)).
In University of London Press Ltd v University Tutorial Press Ltd, there was a contract that required assignment of copyright. "The examiner was the first owner, and he has not assigned the copyright in writing signed by him or his agent. The copyright therefore remains in the examiners, subject to the obligation under the contract of employment to assign it to the University or as it may direct." Even though copyright was owed to the University, it didn't become theirs until the written assignment took place.
Similarly, in the US, "Copyright in a work protected under this title vests initially in the author or authors of the work.", and "in the case of a work made for hire, the employer or other person for whom the work was prepared is considered the author" (17 U.S.C § 201).
"A transfer of copyright ownership, other than by operation of law, is not valid unless an instrument of conveyance, or a note or memorandum of the transfer, is in writing and signed by the owner of the rights conveyed or such owner's duly authorized agent." (17 U.S.C § 204)
In both Canada and the US, transfer of copyright requires an affirmative act (a written, signed document, or a note or memorandum, etc.)
Since nothing in your description indicates that the author has prepared a signed, written, instrument of conveyance or note or memorandum of the transfer of copyright, the copyright still belongs to the original author.